Asiana Flight 214’s pilots caused the crash last year of their airliner carrying more than 300 people by bungling a landing approach in San Francisco, including inadvertently deactivating the plane’s key control for airspeed, the National Transportation Safety Board concluded Tuesday.

But the board also said the complexity of the Boeing 777’s autothrottle and auto flight director — two of the plane’s key systems for controlling flight — contributed to the accident. The board also faulted materials provided to airlines by Boeing that fail to make clear under what conditions the autothrottle doesn’t maintain speed.

The 777 has been in service 18 years and is one of the world’s most popular wide-bodied airliners, especially for international travel. Until last year’s accident, it had not been involved in a single fatal crash.

The board’s acting chairman, Chris Hart, warned that the accident underscores a problem that has long troubled aviation regulators around the globe — that increasingly complicated automated aircraft controls designed to improve safety are also creating new opportunities for error.

The Asiana flight crew “over-relied on automated systems that they did not fully understand,” Hart said.

“In their efforts to compensate for the unreliability of human performance, the designers of automated control systems have unwittingly created opportunities for new error types that can be even more serious than those they were seeking to avoid,” he said.

The South Korea-based airline’s pilot training also was faulted.

Of the 307 people on board Flight 214, three Chinese teens were killed in the July 6, 2013 crash. Nearly 200 were injured, including 49 seriously. It remains the only fatal passenger airline crash in the U.S. in the past five years.

Asiana Airlines said it has already implemented the NTSB’s training recommendations, and that it agreed with the NTSB’s finding that one factor was the complexity of the autothrottle and autopilot systems, as well as their descriptions in Boeing training manuals.

Boeing immediately rejected the notion that the 777’s automated systems contributed to the accident, pointing to the aircraft’s safety record.

“The auto-flight system has been used successfully for over 200 million flight hours across several airplane models, and for more than 55 million safe landings,” the company said in a statement. “The evidence collected during this investigation demonstrates that all of the airplane’s systems performed as designed.”

The safety board “did the right thing,” said Ilyas Akbari, a Los Angeles attorney representing 16 of Flight 214’s passengers. “It took courage to call out Boeing because it’s an American manufacturer” and one of the nation’s largest employers and exporters.

But most of the fault lies with Asiana and its pilots, Akbari said.

Investigators said the flight’s three veteran pilots made 20 to 30 different errors, some minor and others significant, during the landing approach on July 6, 2013.

Two of the three teens who were killed weren’t wearing their seatbelts and were thrown from the plane. The third teen was hit on the head by one of the plane’s doors but survived the crash. She was killed when, lying unconscious, she was run over by two San Francisco Fire Department vehicles in the chaos afterward. Nearly 200 people were injured.

In a statement, Fire Chief Joanne Hayes-White said Tuesday they’ve already enhanced their training and that she is “extremely proud” of all the first responders.